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But lack of reduced carbon can limit the growth of bacteria a great deal.
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Yes that was one thing I thought about when I read the article. It was like all DOC always was bad, when it isn't. Too much is bad though.
Knowing how much is bad comes with experience:
I have completely stable CO2-injection (1 bubble/sec, never measures pH) and the same PMDD-dosage each week. If the pearling from plants diminishes I know there is nothing wrong with light, CO2 or nutrients. It's to much mulm/DOC/POC, and most of the time it's inside the canister. Rinsing it usually brings back the pearling instantaneously.